Butterflying and Birding in Northern Greece

We took a guided trip to Northern Greece in June 2024. It was a butterflying and birding holiday, although there were also plenty of moths, the odd lizard and a jackal. I took a lot of photos but have finally sorted and processed the butterflies. It was too hot and dry much of the time, but we still managed to see at least 70 species of the possible 120 or so that are present in the region. Compare this to the mere 60 we have in the UK, many of which are rarely seen.

Balkan Copper, Lycaena candens
Balkan Copper, Lycaena candens
Balkan Marbled White, Melanargia larissa
Balkan Marbled White, Melanargia larissa
Balkan Marbled White, Melanargia larissa
Balkan Marbled White, Melanargia larissa
Clouded Yellow, Colias croceus
Clouded Yellow, Colias croceus
Clouded Yellow, Colias croceus
Clouded Yellow, Colias croceus
Clouded Yellow, Colias croceus
Clouded Yellow, Colias croceus
Eastern Baton Blue, Pseudophilotes vicrama
Eastern Baton Blue, Pseudophilotes vicrama
Ilex Hairstreak, Satyrium ilicis
Ilex Hairstreak, Satyrium ilicis
Lang's Short-tailed Blue, Leptotes pirithous
Lang’s Short-tailed Blue, Leptotes pirithous
Lang's Short-tailed Blue, Leptotes pirithous
Lang’s Short-tailed Blue, Leptotes pirithous
Little Tiger Blue, Tarucus balkanicus, aka Balkan Pierrot
Little Tiger Blue, Tarucus balkanicus, aka Balkan Pierrot
Little Tiger Blue, Tarucus balkanicus, aka Balkan Pierrot
Little Tiger Blue, Tarucus balkanicus, aka Balkan Pierrot
Mallow Skipper, Carcharodus alceae
Mallow Skipper, Carcharodus alceae
Queen of Spain Fritillary, Issoria lathonia
Queen of Spain Fritillary, Issoria lathonia
Queen of Spain Fritillary, Issoria lathonia
Queen of Spain Fritillary, Issoria lathonia
Queen of Spain Fritillary, Issoria lathonia
Queen of Spain Fritillary, Issoria lathonia
Scarce Swallowtail, Iphiclides podalirius, aka Sail Swallowtail
Scarce Swallowtail, Iphiclides podalirius, aka Sail Swallowtail
Sooty Copper, Lycaena tityrus
Sooty Copper, Lycaena tityrus
Sooty Copper, Lycaena tityrus
Sooty Copper, Lycaena tityrus
Wall Brown, Lasiommata megera
Wall Brown, Lasiommata megera
Nettle-tree Butterfly, Libythea celtis, aka European Beak
Nettle-tree Butterfly, Libythea celtis, aka European Beak

Some of the butterflies we saw in Greece. Of this small collection, only the Wall Brown and the Clouded Yellow are seen in the UK. The latter usually only as a rare migrant or occasionally in larger numbers in a so-called irruption year.

Balkan Copper, Lycaena candens
Balkan Marbled White, Melanargia larissa
Clouded Yellow, Colias croceus
Eastern Baton Blue, Pseudophilotes vicrama
Ilex Hairstreak, Satyrium ilicis
Lang’s Short-tailed Blue, Leptotes pirithous
Little Tiger Blue, Tarucus balkanicus, aka Balkan Pierrot
Mallow Skipper, Carcharodus alceae
Queen of Spain Fritillary, Issoria lathonia
Scarce Swallowtail, Iphiclides podalirius, aka Sail Swallowtail
Sooty Copper, Lycaena tityrus
Wall Brown, Lasiommata megera
Nettle-tree Butterfly, Libythea celtis, aka European Beak

Butterflies of Greece Batch 123456789

Moths matter in Macedonia

I’ve sorted through my moth photos from our recent Birding and Butterflying trip to Greece with Greenwings. I’m sure I saw a few more, but this is the list of 43 compiled from all my photos. NFM is “New for me”.

Common Name Scientific Name NFM
Black-veined Moth Siona lineata Y
Bright Wave Idaea ochrata Y
Broad-bordered Bee Hawk-moth Hemaris fuciformis Y
Brown China-mark Elophila nymphaeata Y
Burnet Companion Euclidia glyphica N
Catocala nymphaea Catocala nymphaea Y
Chrysocrambus cassentiniellus Chrysocrambus cassentiniellus Y
Common Heath Ematurga atomaria N
Crambus perlella Crambus perlella N
Crocidosema plebejana Crocidosema plebejana Y
Dingy Shell Euchoeca nebulata Y
Chequered Wave Idaea moniliata Y
Dotted Border Wave Idaea sylvestraria Y
Euclasta splendidalis Euclasta splendidalis Y
Eucosma cana Eucosma cana N
Euplocamus ophisa Euplocamus ophisa Y
Five-spot Burnet Zygaena trifolii N
Forester Adscita statices Y
The Geometrician Grammodes stolida Y
Grass Rivulet Perizoma albulata Y
Gypsy Moth Lymantria dispar N
The Handmaid Dysauxes ancilla Y
Hummingbird Hawk-moth Macroglossum stellatarum N
Knot Grass Acronicta rumicis N
Lead Belle Scotopteryx mucronata Y
Mother Shipton Callistege mi N
Nine-spotted Moth Amata phegea Y
The Passenger Dysgonia algira Y
Poplar Grey Acronicta megacephala N
Purple-barred Yellow Lythria cruentaria Y
Pyrausta purpuralis Pyrausta purpuralis N
Ragades pruni Ragades pruni Y
Riband Wave Idaea aversata N
Rusty Dot Pearl Udea ferrugalis N
Silver Ground Carpet Xanthorhoe montanata Y
Six-spot Burnet Zygaena filipendulae N
Slender Scotch Burnet Zygaena loti Y
Small Fan-footed Wave Idaea biselata Y
Speckled Yellow Pseudopanthera macularia Y
Spotted Sulfur Emmelia trabealis Y
Spurge Hawk-moth Hyles euphorbiae Y
Transparent Burnet Zygaena purpuralis Y
Treble-bar Aplocera plagiata N
Wood Tiger Arctia plantaginis Y

We were in Northern Greece near Lake Kerkini, staying in a village called Chrisochorafa and trekking the mountains and lakes, of what was known as Macedonia in ancient times. We were touched the border of present-day North Macedonia at one point and on another day were not too far from Bulgaria.

Moths of Macedonia

I recorded some 43 moths on our trip to Greece this year, and perhaps saw a few more micros of which I didn’t get photos. I’ve tabulated the full list as it stands in a separate post entitled Moths Matter in Macedonia.

As well chasing the obvious on our Greenwings butterflying and birding trip to Greece, there was also time for a few moths. Some during the day, some that overnighted in the stairwell of our little hotel, and some from fellow amateur Lepidopterist, Martine’s field trips around the village of Chrisochorafa. Thanks to Martine and Michael for IDs on the local species and for Mrs Sciencebase’s eagle eyes in spotting much of the wildlife.

Pyropteron minianiformis
Pyropteron minianiformis

Spotted my one and only clearwing moth of the trip, Pyropteron minianiformis, on our first day trekking through the disused Vironia Quarry. This was also where we saw our first Little Tiger Blue, Ilex Hairstreak, Clouded Yellow, and Nettle-tree butterflies, Masked Shrike, Levant Sparrowhawk and overhead Ravens too.

Closeup of one of the tiniest, but perhaps showiest, moths we saw in Greece. This is a Balkan species known as Euplocamus ophisa (Tineidae), which feeds on bracket fungi and rotten wood. The male's antennae are almost as long its wings!
Closeup of one of the tiniest, but perhaps showiest, moths we saw in Greece. This is a Balkan species known as Euplocamus ophisa (Tineidae), which feeds on bracket fungi and rotten wood. The male’s antennae are almost as long its wings!

We also saw our first tortoise of the trip. We had seen our first Bee-eaters, White Stork, and Honey Buzzard en route, and heard our first frogs. There will be more about the birds and other wildlife in a subsequent post, once I have processed those photos.

Euplocamus ophisa
Euplocamus ophisa

The yellow and black micro moth with large, feathery antennae is a male Euplocamus ophisa (Tineidae). Spotted several of these in the stream-side woodland where the Glider butterflies were seen. No ID for the arachnid.

Speckled Yellow, Pseudopanthera macularia
Speckled Yellow, Pseudopanthera macularia

Mrs Sciencebase spotted the first Speckled Yellow (Pseudopanthera macularia, Geometridae) of the trip, at the Lailias Ski Centre, I think it was.

Catocala nymphaea
Catocala nymphaea

I spotted this large moth, Catocala nymphaea (Erebidae), which was roosting on a derelict building near a barely used dirt track we butterflied. It is related to Red Underwing, Crimson Underwing, Clifden Nonpareil, but, while it has orange on its hindwings, it is not an orange underwing.

Grammodes stolida
The Geometrician, Grammodes stolida

The Geometrician, Grammodes stolida (Erebidae). Potted by Martine on her late-night field trip not far from Chrisochorafa.

Acronicta rumicis
Knot-grass, Acronicta rumicis, larva

Knot-grass (Acronicta rumicis, Noctuidae) larva feeding on spurge, I believe, Vironia Quarry

Idaea aversata
Riband Wave, Idaea aversata

The filled band on this Riband Wave, Idaea aversata (Geometridae), is rather more pink than I have seen before.

Lead Belle, Scotopteryx mucronata
Lead Belle, Scotopteryx mucronata
Hemaris fuciformis
Broad-bordered Bee Hawk-moth, Hemaris fuciformis
Hemaris fuciformis
Broad-bordered Bee Hawk-moth, Hemaris fuciformis

We were watching lots of butterflies and the nectaring Hummingbird Hawk-moths on a purple patch of vetch at the side of the road up towards the Lailias Ski Centre when a couple of Broad-bordered Bee Hawk-moth, (Hemaris fuciformis, Sphingidae) put in a timely appearance.

Hyles euphorbiae larvae
Spurge Hawk-moth (Hyles euphorbiae, Sphingidae, larvae

Michael spotted a couple of Spurge Hawk-moth (Hyles euphorbiae, Sphingidae) larvae feeding on some roadside spurge, like they do.

Euclasta splendidalis
Euclasta splendidalis

Early-morning phone snap of a long-legged moth which was roosting at dawn at our accommodation. Euclasta splendidalis (Crambidae) is restricted to SE Europe onwards to Turkey and the Middle East. The food plant is a vine called Periploca graeca (Asclepiadaceae – Milkweed family).

Amata phegea
Nine-spotted Moth, Amata phegea

There were lots of Nine-spotted Moth around on our wanderings. They are also known as the Yellow-belted Burnet (Amata phegea, Erebidae) or in Michael’s parlance, the “lazy moth” on account of their general indolence.

Wood Tiger, Parasemia plantaginis
Wood Tiger, Parasemia plantaginis

Wood Tiger (Parasemia plantaginis, Erebidae) spotted and potted by Martine before being released back on site for a photo or two.

The Passenger, Dysgonia algira
The Passenger, Dysgonia algira

The Passenger (Dysgonia algira, Noctuidae) was fluttering around the hotel stairwell with several micros when we returned from our evening meal with the Little Owls.

Black-veined Moth, Siona lineata
Black-veined Moth, Siona lineata

Black-veined Moth (Siona lineata, Geometridae), not to be confused with the Black-veined White butterfly, which we also saw.

Hummingbird Hawk-moth, Macroglossum stellatarum
Hummingbird Hawk-moth, Macroglossum stellatarum
Bright Wave, Idaea ochrata
Bright Wave, Idaea ochrata
The Forester being consumed by an arachnid
The Forester being consumed by an arachnid

The Handmaid

The Handmaid

Treble-bar
Treble-bar
Dotted Border Wave
Dotted Border Wave
Slender Scotch Burnet
Slender Scotch Burnet
Six-spot Burnet
Six-spot Burnet
Transparent Burnet
Transparent Burnet
Rhagades pruni
Rhagades pruni
Purple-barred Yellow
Purple-barred Yellow
Common Heath
Common Heath
Mother Shipton
Mother Shipton
Chrysocrambus cassentiniellus
Chrysocrambus cassentiniellus
Small Fan-footed Wave, Idaea biselata
Small Fan-footed Wave, Idaea biselata – Not 100% on the idea
Pyrausta purpuralis
Pyrausta purpuralis
Female Gypsy Moth
Female Gypsy Moth, phone snap
Dotted Sulphur
Spotted Sulphur, phone snap
Brown China-mark, Elophila nymphaeata
Brown China-mark, Elophila nymphaeata (wobbly shot on the boat, too big a lens). Several on the boat, the larvae feed on aquatic plants.
Not a great photo of Chequered Wave
Not a great photo of Chequered Wave

The Macedonia of my blog headline refers to the ancient Greek region which coincides with the area of modern, northern Greece where we were staying and exploring. North Macedonia was west of us, just over the border. At one point, we were within a few metres of the North Macedonian border patrol as we headed for a known site – the woodland and scrub around the Doiran Memorial – seeking, unsuccessfully as it turned out, the Tessellated Skipper butterfly.

The Petaloúda Paparazzi

Mrs Sciencebase and I spent a week in the beautiful wilds of northern Greece not far from the Bulgarian and North Macedonian borders in early June. At first glance, it was a holiday, but it was more realistically a fascinating international biology field trip with lots of butterflies, moths, birds, and plants to photograph and record. The list of butterflies we saw and recorded can be found below.

Nemoptera sinuata, one of the spoonwinged lacewings
Nemoptera sinuata, not a butterfly, not a moth, not a bird, a spoonwinged lacewing

We had 6:30am starts most days except the 5am on Lake Kerkini boat-trip day, expertly piloted by Niko. It was certainly not the usual lounging-by-the-pool-all-day type holiday with lingering lie-ins et cetera. There was, however, all those butterflies and birds. Delicious Greek food and quaffable Greek beer every evening certainly felt holiday-like!

Mrs Sciencebase in Dora the Explorer mode, photographing a chicory plant
Mrs Sciencebase in Dora the Explorer mode, photographing a chicory plant

We were staying in a small town called Chrisochorafa, which isn’t far from the beautiful Lake Kerkini with its hundreds of pelicans (Dalmation and Great White), tens of thousands of Great Cormorant, numerous Spoonbill, various heron and egret species, and on and on!

Our balcony view of the "twin towers" in Chrysochorafa - church and water
Our balcony view of the “twin towers” in Chrysochorafa – church and water. Chrysochorafa, meaning “golden fields”.

Our expert eco-guide, Michael, kindly drove us all over the region. We swung way west to the border with North Macedonia, but also east and north almost to Bulgaria too. Michael, is a diptera and lepidoptera expert and did a bit of independent flycatching in between identifying butterflies and birds for his research. He hopes to identify the groups that may well have been lost to regions ravaged by forest fires in northeastern Greece in recent years.

Our expert guide, driver, and now good friend, Michael
Our expert guide, driver, and now good friend, Michael “D”

We also had one other travelling companion, Martine – a modestly expert amateur lepidopterist who was a great help to us in spotting and identifying the novel species of Lepidoptera. We hiked several km most days even when it was sometimes 38 degrees Celsius in the shade.

Martine, butterflying, coffee in hand
Martine, butterflying, coffee in hand

Somehow, I managed to take more than 11000 photos. Too much of the time I had my camera in rapid-burst mode. I’ve plucked more than 500 from the SD cards. I will pull out and process what I think are the best to share here on the Sciencebase blog and on my social media. I may be some time. You Have Been Warned.

European Green Lizard
European Green Lizard

I now have photographs of dozens of species of butterfly that I’d not seen before, and we saw well over 73 species in total. Lots of birds and more than 40 moth species, more than a couple of dozen of which aren’t present in the UK and I’d not seen before. 90+ birds seen and/or heard.

Balkan Copper butterfly, Lycaena candens, photographed on the ski slope at Lailias near Serres in Northern Greece
Balkan Copper butterfly, Lycaena candens

I didn’t do a lot of handheld focus stacking, but I did try with the above Balkan Copper photo. It was on the ski slope at Lailias, no snow, obviously. There were quite a few other butterflies and day-flying moths here.

Wild Strawberry
Wild Strawberry

It’s worth noting that contractors were strimming the lovely meadow at the foot of the ski slope for some unknown reason. This will inevitably have destroyed thousands of blooms, sending the invertebrates that were thriving there into oblivion. There was presumably a good reason for the strimming, but it put paid to our butterflying on that patch.

Bladetail dragonfly, Lindenia tetraphylla with prey
Bladetail dragonfly, Lindenia tetraphylla with prey

Overall, we had a fabulous trip although some of our target species were entirely absent from even the most likely sites. Tessellated Skipper, for instance. Indeed, its larval food plant was absent from the site too. Temperatures were too high for June in this region for the whole week. This may well have contributed to the much lower than anticipated numbers and diversity of Lepidoptera, although we still saw plenty.

Common Awl Robber Fly, Neoitamus cyanurus
A bristly fly of dry, grassland, ID to be confirmed
Lion detail from The Doiran Memorial, a Commonwealth War Graves Commission war memorial near Doirani in northern Greece not far from the present North Macodian border
Lion detail from The Doiran Memorial

Praying mantis, Empusa fasciataTwo Praying Mantis, Empusa fasciata

Small Pincertail, Onychogomphus forcipatus, Green-eyed Hooktail, Green-eyed Hook-tailed Dragonfly
Small Pincertail, Onychogomphus forcipatus

Butterflies of Greece Batch 123456789


Beautiful Demoiselle, Calopteryx virgo, presumably the Balkan "festiva" sub-species
Beautiful Demoiselle, Calopteryx virgo

Great Green Bush Cricket, Tettigonia viridissimaGreat Green Bush Cricket, Tettigonia viridissima

Schmidt's Marbled Bush Cricket, Eupholidoptera schmidti
Schmidt’s Marbled Bush Cricket, Eupholidoptera schmidti
Balkan Pond Turtle, Balkan Terrapin or Western Caspian Terrapin, Mauremys rivulata
Balkan Pond Turtle, Balkan Terrapin or Western Caspian Terrapin – Mauremys rivulata
Balkan Frog, Pelophylax kurtmuelleri
Balkan Frog, Pelophylax kurtmuelleri
There were unseeing eyes watching the couple in their hammock
There were unseeing eyes watching the couple in their low-slung hammock
Eastern Green Lizard
Why isn’t the Eastern Green Lizard called the Balkan Blue-throated Lizard?
White-faced Bush-Cricket, Decticus albifrons
White-faced Bush-Cricket, Decticus albifrons
Speckled Bush-cricket, Leptophyes punctatissima
Speckled Bush-cricket, Leptophyes punctatissima
Golden-bloomed Longhorn Beetle, Agapanthia villosoviridescens, in cop
Golden-bloomed Longhorn Beetle, Agapanthia villosoviridescens, in cop
Robber Fly, Neoitamus cyanurus
Robber Fly, Neoitamus cyanurus or a close relative
Black-striped Longhorn Beetle, Stenurella melanura, in cop
Black-striped Longhorn Beetle, Stenurella melanura, in cop
Hotel Limnaio, Chrysochórafa, northern Greece (Central Macedonia)
Hotel Limnaio, Chrysochórafa, northern Greece (Central Macedonia)
Hermann's Tortoise, Testudo hermanni
Hermann’s Tortoise, Testudo hermanni
Spotted Orbweaver sp
Spotted Orbweaver sp
Red Shield-bug, Carpocoris mediterraneus, in cop
Red Shield-bug, Carpocoris mediterraneus, in cop
Mediterranean house gecko, Hemidactylus turcicus
Mediterranean house gecko, Hemidactylus turcicus
Grape Wood Borer, Chlorophorus varius
Grape Wood Borer, Chlorophorus varius
The Minibus
The Minibus
Bridge under which I think I saw a Camberwell Beauty
Bridge under which I think I saw a Camberwell Beauty
The Azure and White
The Azure and White – national flag of Greece against the dusk

June 2024 Greek butterfly tick-list

Here is the complete list of butterfly species of which I have now published photographs on Sciencebase. There were several others that we saw that I didn’t get photos of or that didn’t warrant sharing – Essex Skipper, Small Skipper, Dark Green Fritillary etc. I think the total count for the trip for me and Mrs Sciencebase was 73+ butterfly species.

  1. Blue, Amanda’s (Polyommatus amandus)
  2. Blue, Eastern Baton (Pseudophilotes vicrama)
  3. Blue, European Common (Polyommatus icarus)
  4. Blue, Green-underside (Glaucopsyche alexis)
  5. Blue, Iolas (Iolana iolas)
  6. Blue, Lang’s Short-tailed (Leptotes pirithous)
  7. Blue, Large (Phengaris arion)
  8. Blue, Little Tiger (Tarucus balkanicus)
  9. Blue, Mazarine (Cyaniris semiargus,)
  10. Blue, Small (Cupido minimus)
  11. Brown Argus (Aricia agestis)
  12. Brown, Large Wall (Lasiommata maera)
  13. Brown, Lattice (Kirinia roxelana)
  14. Brown, Meadow (Maniola jurtina)
  15. Brown, Northern Wall (Lasiommata petropolitana)
  16. Brown, Wall (Lasiommata megera)
  17. Cardinal (Argynnis Pandora)
  18. Clouded Apollo (Parnassius mnemosyne)
  19. Clouded Yellow (Colias croceus)
  20. Common Yellow Swallowtail (Papilio machaon)
  21. Copper, Balkan (Lycaena candens)
  22. Copper, Lesser Fiery (Lycaena thersamon)
  23. Copper, Purple-shot (Lycaena alciphron)
  24. Copper, Sooty (Lycaena tityrus)
  25. Eastern Festoon (Allancastria cerisyi)
  26. Fritillary, Heath (Melitaea athalia)
  27. Fritillary, Knapweed (Melitaea phoebe)
  28. Fritillary, Lesser Marbled (Brenthis ino)
  29. Fritillary, Lesser Spotted (Melitaea trivia)
  30. Fritillary, Marbled (Brenthis daphne)
  31. Fritillary, Niobe (Fabriciana niobe)
  32. Fritillary, Pearl-bordered (Boloria euphrosyne)
  33. Fritillary, Queen of Spain (Issoria lathonia)
  34. Fritillary, Silver-washed (Argynnis paphia)
  35. Fritillary, Spotted (Melitaea didyma)
  36. Grayling, Eastern Rock (Hipparchia syriaca)
  37. Grayling, Great Banded (Brintesia circe)
  38. Hairstreak, Blue-spot (Satyrium spini)
  39. Hairstreak, Green (Callophrys rubi)
  40. Hairstreak, Ilex (Satyrium ilicis)
  41. Hairstreak, Sloe (Satyrium acacia)
  42. Hairstreak, White-letter (Satyrium w-album)
  43. Heath, Pearly (Coenonympha arcania)
  44. Heath, Small (Coenonympha pamphilus)
  45. Hermit (Chazara briseis)
  46. Nettle-tree Butterfly (Libythea celtis)
  47. Painted Lady (Vanessa cardui)
  48. Scarce Swallowtail (Iphiclides podalirius)
  49. Skipper, Dingy (Erynnis tages)
  50. Skipper, Grizzled (Pyrgus malvae)
  51. Skipper, Large (Ochlodes sylvanus)
  52. Skipper, Mallow (Carcharodus alceae)
  53. Skipper, Yellow-banded (Pyrgus sidae)
  54. Southern White Admiral (Limenitis reducta)
  55. Tortoiseshell, Large (Nymphalis polychloros)
  56. Tortoiseshell, Small (Aglais urticae)
  57. White, Balkan Marbled (Melanargia larissa)
  58. White, Black-veined (Aporia crataegi)
  59. White, Eastern Bath (Pontia edusa)
  60. White, Marbled (Melanargia galathea)
  61. White, Small (Pieris rapae)
  62. White, Wood (Leptidea sinapis)
  63. Woodland Ringlet (Erebia medusa)

Technical footnotes

I took rather a lot of photos on this week-long trip (well over 10,000) and then spent the weekend pulling out what I think are the best. I’ve got about 500 that I need to work through and do a second-line purge.

I will process the select few from the camera RAW files using DxO PureRaw4, which I’ve mentioned previously. This software carries out corrections to the photos you take based on the known distortions associated with your camera and lens combination. Its main purpose however is to remove noise from an image without removing detail. It does this very well, improving any photo you feed it to the tune of 3 or so full stops of ISO. This can make all the difference with photos shot in the limited light of a woodland or at twilight.

Some of the photos need a different kind of processing and I’ll use Topaz Sharpen for the ones that could do with a bit more tightening up to remove motion blur, for instance.